In the majority of bird species synchronous hatching is the norm.
This is achieved simply: the parent does not commence incubation until
the clutch of eggs, however large, is complete.
Most birds will lay an egg each day until an appropriate
trigger indicates that the clutch is complete. This is why I can collect
an egg a day from each of my hens - they keep laying until incubation
is triggered by some stimulus. This could be the feel of a full clutch
against the brood patch on a bird's belly, but there is also some
endogenous control factor. For example, one of my hens may turn "broody"
- she will cease laying and commence incubation on only one egg if that
is all I have left her with.
In some other groups of birds - notably owls, raptors and
cormorants - incubation commences when the first egg is laid, leading
to sequential hatching, with the first chick gaining a significant
advantage over later ones. This is very noticeable in barn owls, where
five or six eggs laid at daily intervals lead to the oldest chicks being
almost a week older than the youngest. This strategy ensures maximum
chick survival in species with an unpredictable food source: in years of
plenty all the offspring get enough food, but in years when food is
less readily available the oldest, largest chicks survive and dominate
the younger ones - which almost inevitably perish and may be eaten by
their siblings.
This strategy sees its most elegant expression in the Cain-and-Abel syndrome, which is manifested particularly well in eagles.
The first egg hatches two to three days before the second and, when
food is scarce, there seems to be a degree of inevitability in the way
the older chick persecutes its younger sibling to the point of death.
Norman McCanch, Canterbury, Kent, UK